Hard
Derivatives of Epidermis.
B.
Sc- III Zoology. Dr. Kamble V. S.
- Following types of horns are known:
- 1.
Hollow Horns
- 2.
Pronghorn
- 3.
Antlers
- 4.
Hair Horn
- 5.
Giraffe Horns
- Hollow Horns
Horns are found in
ungulate (even-toed hoofed) mammals only. True horns of the hollow type are
found in pronghorn, cattle, antelope, sheep and goats consist of an inner core
of bone which is an outgrowth from the frontal bone. It is encased in a
keratinised, epidermal covering. True horns continuously grow throughout life
and are not shed.
Pronghorn
Antlers
Antlers are found in
the males of deer family, but they are present in both sexes in reindeer and
caribou. An antler consists of a branching solid outgrowth of the frontal bone
formed of dense connective tissue. It is covered during growth by hairy,
vascular skin called ‘velvet’. The velvet is shed exposing the antler naked
when the antler reaches full growth.
Thus, the antler
consists only of dermal bone. The bony antler is also shed annually after the
breeding season, and a new antler develops. Antlers are solid mesodermal bone,
but they are formed under the influence of the integument. Formation of antlers
is controlled by the hormones of testes and anterior lobe of the pituitary.
Rhinoceros or Hair Horn
horn has no skeletal element. It is made by keratinised cells of the epidermis and consists of matted keratin fibres bound together, but its fibres are not true hair. It is a permanent epidermal structure and if broken it grows again. There is one horn in the Indian rhinoceros and two in the African species.
Giraffe Horns:
They develop from
cartilaginous protrusions which are present at birth. They ossify and fuse at
the top of the skull, where they appear as knobs permanently covered with
living skin and hair. Giraffe possesses three of these knobs, one is median and
anterior to the other two. These horns are short, unbranched and are permanent,
and are present in both sexes.
Digital tips
In amniota the distal
ends of digits have claws, nails or hoofs formed from the horny layer of the
epidermis. They grow parallel to the surface of the skin and are built on the
same plan.
Claws:
Claws
made their appearance first in the reptiles. A claw is made of a hard horny
dorsal scale-like plate called unguis and a relatively soft ventral subunguis,
both converge terminally and cover the terminal part of the last phalanx. Claws
of reptiles and birds are similar but in mammals the subunguis is much reduced
and is continuous with a pad at the end of a digit. In cat family claws are
retractile.
ii.
Nails:
They
are found in primates. The dorsal unguis is large and flat and subunguis is
soft and much reduced. Tip of the digit forms a sensitive and vascular pad over
which the nail groove is present. It is formed by the invagination of
epidermis. Growth of the unguis takes place from the nail root lying below the
skin in the nail groove.
iii.
Hoofs:
They
are found in ungulates. The horny unguis is thick and around the end of the
digit, and encloses the thickened subunguis which touches the ground. Subunguis
surrounds the soft, horny cuneus. Tip of digit, thus, forms a pad containing a
blunt phalanx. Nails and hoofs of mammals are modified claws. Whalebone plates
of toothless whales are also the modification of stratum corneum.
Hair:
Hair is found only in mammals. It projects at an acute angle from the skin. Hair covers the entire integument in most (furred mammals), but in others only traces are left, such as whales have only a few core hairs on the snout.
But
during development the body of the embryos of all mammals is covered with a
coating of fine hair called lanugo which is usually shed before birth and
replaced by a new one. Hair is entirely epidermal in origin.
Hairs
are not modified scales but are new outgrowths of the epidermis only. A hair
has an upper projecting shaft and a lower root lying in a hair follicle which
is a sunken pit in the dermis. The shaft is made of only dead, keratinised
cells. The part of the hair protruding above the skin is dead.
The
hair shaft has an external cuticle of transparent overlapping cells which have
lost their nuclei, inside the cuticle is a cortex (middle part of hair)
containing shrivelled cells and pigments, and a central core or medulla having
air spaces.
Feathers in Bird
Feathers
are found only in birds and are formed from the epidermis in which the stratum
corneum is highly specialised. Feathers are light, strong, elastic, waterproof
and show many colours due to pigments and structural arrangement. The pigments
are carotenoids and melanins. Carotenoids are frequently called lipochromes
which are soluble in fat solvents like methanol, ether or carbon disulphide,
and insoluble in water.
A typical feather consists of following parts.
1.
Central axis or
Scapus
2.
Vexillum or
vane.
3.
- 1.
Basal calamus
- 2. Upper shaft or rachis.